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"On my mark, you, Bolton and Levine are gonna break formation and head back to the city -- take out anything that we missed."
John Sheppard[src]

Bolton was a Tau'ri and member of the Atlantis expedition. In 2007 when Atlantis left Lantea, Bolton was one of the Puddle Jumper pilots who destroyed the asteroids blocking Atlantis's path, which indicated he has the ATA gene. (SGA: "Adrift")